Entry 2 senses Webster, 1913 Philomela /fĭlˌ-əm-ēl'-ə/ · Phil·o·me·la · IPA /fˌɪləmˈiːlə/ 01 n. The nightingale; philomel. 1. The nightingale; philomel. 2. A genus of birds including the nightingales.(Zool.)